Scientific publications

Up to date as of 10 October 2020. For the full text of items not available for download here, or hardcopy versions, please contact me. For estimated citation indexes and links to citing papers, please consult my Google Scholar profile, and for broader impacts of my peer-reviewed research, see my Impactstory profile; I also maintain a profile on Mendeley. Because I consider public outreach an important part of doing science, I frequently discuss my own projects on popular websites to which I contribute. Links to relevant online articles follow the listings for papers I’ve covered in that format.

Jump to my peer-reviewed articles, other scholarly publications, selected presentations, or other writing.

Peer-reviewed articles

Top | Peer-reviewed articles | Other publications | Selected presentations

Yoder JB, G Gomez, and CJ Carlson. 2020. Zygomorphic flowers have fewer potential pollinators. Biology Letters. 16(9): 20200307. doi: 10.1098/rsbl.2020.0307. Data on Dryad, doi: 10.5061/dryad.gxd2547j3. Full-text preprint on bioRxiv, doi: 10.1101/743872.

Briscoe Runquist RD, AJ Gorton, JB Yoder, NJ Deacon, JJ Grossman, Shan Kothari, MP Lyons, SN Seth, P Tiffin, and DA Moeller. 2020. Context dependence of local adaptation to abiotic and biotic environments: A quantitative and qualitative synthesis. American Naturalist.195(3): 412-431. doi: 10.1086/707322. Full text (PDF, 2MB) — Featured in a PNAS Journal Club post.

Mahony CR, IR MacLachlan, BM Lind, JB Yoder, T Wang, SN Aitken. 2019. Evaluating genomic data for management of local adaptation in a changing climate: A lodgepole pine case study. Evolutionary Applications. 13(1):116-131. doi: 10.1111/eva.12871. Full text (PDF, 2MB)

Terrill RS JM Maley, WLE Tsai, KB Fistanic, RJ Freeland, A Franceschelli, B Lewis-Smith, LL Lu, and JB Yoder. 2019. Tricolored blackbirds feeding in Joshua tree inflorescences. Western Birds. 50: 180-182. doi: 10.21199/WB50.3.6. Full text (open access).

Mattheis A, D Cruz-Ramírez De Arellano, and JB Yoder. 2019. A model of Queer STEM identity in the workplace. Journal of Homosexuality, doi: 10.1080/00918369.2019.1610632. Full text (PDF, 1.9MB)

Yoder JB and P Tiffin. 2017. Sanctions, partner recognition, and variation in mutualism. American Naturalist. 190(4): 491-505. doi: 10.1086/693472. Data on Dryad, doi: 10.5061/dryad.p2s02. Full-text preprint on bioRxiv, doi: 10.1101/049593.

Yoder JB and P Tiffin. 2017. Effects of gene action, time since selection, and marker density on the performance of landscape genomic scans of local adaptation. Journal of Heredity. 109(1): 16-28. doi: 10.1093/jhered/esx042. Full text (PDF, 1.2MB). Data on Dryad, doi: 10.5061/dryad.p12q4.


Yoder JB. 2016. Understanding the coevolutionary dynamics of mutualism with population genomics. American Journal of Botany. 103(10):1742-52. doi: 10.3732/ajb.1600154. Full text (PDF, 661KB). Part of a Special Issue on intimate mutualism, co-edited with J Leekens-Mack.

Yoder JB and A. Mattheis. 2016. Queer in STEM: Workplace experiences reported in a national survey of LGBTQA individuals in science, technology, engineering, and mathematics careers. Journal of Homosexuality. 63(1): 1-27. doi: 10.1080/00918369.2015.1078632. Full text (PDF, 499KB)

Hembry DH, JB Yoder, and KR Goodman. 2014. Coevolution and the diversification of life. American Naturalist. 184(4): 425-38. doi: 10.1086/677928. Full text (PDF, 436KB)

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Yoder JB, J Stanton-Geddes, P Zhou, R Briskine, ND Young, and P Tiffin. 2014. Genomic signature of local adaptation to climate in Medicago truncatula. Genetics. 196(4): 1263-75. doi: 10.1534/genetics.113.159319. Full text (PDF, 3.4MB). See also The Molecular Ecologist: Scanning the genome for local adaptation.

Stanton-Geddes J, JB Yoder, R Briskine, ND Young, and P Tiffin. 2013. Estimating heritability with whole-genome sequence data. Methods in Ecology and Evolution. 4(12): 1151-8. doi: 10.1111/2041-210X.12129. Full text (PDF, 717KB).

Stanton-Geddes J, T Paape, B Epstein, R Briskine, JB Yoder, J Mudge, AK Bharti, AD Farmer, P Zhou, R Denny, GD May, S Erlandson, M Sugawara, MJ Sadowsky, ND Young, P Tiffin. 2013. Candidate genes and genetic architecture of symbiotic and agronomic traits revealed by whole-genome, sequence-based association genetics in Medicago truncatula. PLOS ONE. 8(5): e65688. doi: 10.1371/journal.pone.0065688. Full text (open access).


Yoder JB, CI Smith, DJ Rowley, WKW Godsoe, CS Drummond, and O Pellmyr. 2013. Effects of gene flow on phenotype matching between two varieties of Joshua tree (Yucca brevifolia; Agavaceae) and their pollinators Journal of Evolutionary Biology. 26(6): 1220-33. doi: 10.1111/jeb.12134. Full text (PDF, 595KB). Data on Dryad, doi 10.5061/dryad.369q9. See also Nothing in Biology Makes Sense: One of these moths is not like the other … but does that matter to Joshua trees?

Yoder JB, R Briskine, J Mudge, A Farmer, T Paape, K Steele, G Weiblen, AK Bharti, GD May, P Zhou, N Young, and P Tiffin. 2013. Phylogenetic signal variation in the genomes of Medicago (Fabaceae). Systematic Biology. 62(3): 424-38. doi: 10.1093/sysbio/syt009. Full text (PDF, 1.2MB). Data on Dryad, doi 10.5061/dryad.vp634306. See also Nothing in Biology Makes Sense: How many phylogenies are there in a genome? Lots!

Starr TN, KE Gadek, JB Yoder, R Flatz, and CI Smith. 2013. Asymmetric hybridization and gene flow between Joshua trees (Agavaceae: Yucca) reflects differences in pollinator host specificity. Molecular Ecology. 22(2): 437-49. doi: 10.1111/mec.12124. Full text (PDF, 692KB).

Flatz, R, JB Yoder, E Lee-Barnes, and CI Smith. 2011. Characterization of microsatellite loci in Yucca brevifolia (Agavaceae) and cross-amplification in related species. American Journal of Botany. 98: e67-9. doi: 10.3732/ajb.1000468. Full text (PDF, 464KB).

Godsoe W, JB Yoder, CI Smith, CS Drummond, O Pellmyr. 2010. Absence of phenotype matching in an obligate pollination mutualism. Journal of Evolutionary Biology. 23(12): 2739-46. doi: 10.1111/j.1420-9101.2010.02120.x. Full text (PDF, 332KB). See also Denim and Tweed: Coevolutionary constraints may divide Joshua trees.

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Yoder JB and SL Nuismer. 2010. When does coevolution promote diversification? American Naturalist. 176(6): 802-17. doi: 10.1086/657048. Full text (PDF, 536KB). See also Denim and Tweed: Not all species interactions are (co)evolved equal.

Drummond CS, H-J Xue, JB Yoder, and O Pellmyr. 2010. Host-associated divergence and incipient speciation in the yucca moth Prodoxus coloradensis (Lepidoptera: Prodoxidae) on three species of host plants. Heredity. 105(2): 183-96. doi: 10.1038/hdy.2009.154. Full text (PDF, 907KB).

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Yoder JB, E Clancey, S Des Roches, JM Eastman, L Gentry, WKW Godsoe, T Hagey, D Jochimsen, BP Oswald, J Robertson, BAJ Sarver, JJ Schenk, S Spear, and LJ Harmon. 2010. Ecological opportunity and the origin of adaptive radiations. Journal of Evolutionary Biology. 23(8): 1581-96. doi: 10.1111/j.1420-9101.2010.02029.x. Full text (open access). See also Denim and Tweed: When ecological opportunity knocks, does adaptive radiation answer? and my guest post for the Scientific American blog, Ecological opportunity: The seed of evolutionary change in your backyard–and in your veins.

Yoder JB, CI Smith, and O Pellmyr. How to become a yucca moth: Minimal trait evolution needed to establish the obligate pollination mutualism. Biological Journal of the Linnean Society 100(4): 847-55. doi: 10.1111/j.1095-8312.2010.01478.x. Full text (PDF, 656KB). See also Denim and Tweed: Before they were yucca moths.


Smith CI, CS Drummond, WKW Godsoe, JB Yoder, and O Pellmyr. 2009. Host specificity and reproductive success of yucca moths (Tegeticula spp. Lepidoptera: Prodoxidae) mirror patterns of gene flow between host plant varieties of Joshua tree (Yucca brevifolia: Agavaceae). Molecular Ecology. 18(24): 5218-29. doi: 10.1111/j.1365-294X.2009.04428.x. Full text (PDF, 456KB). See also Denim and Tweed: For yucca moths, does (flower) size matter?

Pellmyr O, JB Yoder, and WK Godsoe. 2009. Prodoxus praedictus, sp.n., a new bogus yucca moth from southern California. Journal of the Lepidopterists’ Society 63(3): 137-140. Full text (PDF, 542KB).

Godsoe WK, E Strand, T Esque, CI Smith, JB Yoder, and O Pelmyr. 2009. Divergence in an obligate mutualism is not explained by divergent climatic factors. New Phytologist 183(3): 589-99. doi: 10.1111/j.1469-8137.2009.02942.x. Full text (PDF, 3.7MB). See also Denim and Tweed: Finding Joshua tree’s niche.


Smith CI, WKW Godsoe, S Tank, JB Yoder, and O Pellmyr. 2008. Distinguishing coevolution from covicariance in an obligate pollination mutualism: Asynchronous divergence in Joshua tree and its pollinators. Evolution 62(10): 2672-87. doi: 10.1111/j.1558-5646.2008.00500.x. Full text (PDF, 438KB). See also Denim and Tweed: Joshua tree genetics suggest coevolutionary divergence.

Godsoe W, JB Yoder, CI Smith, and O Pellmyr. 2008. Coevolution and divergence in the Joshua tree/yucca moth mutualism. American Naturalist 171(6): 816-23. doi: 10.1086/587757. Full text (PDF, 447KB).

Gomulkiewicz R, DM Drown, MF Dybdahl, W Godsoe, SL Nuismer, KM Pepin, BJ Ridenhour, CI Smith, and JB Yoder. 2007. Do’s and Don’ts of testing the geographic mosaic theory of coevolution. Heredity 98(5): 249-58. doi: 10.1038/sj.hdy.6800949. Full text (PDF, 311KB).

Other scholarly works

Top | Peer-reviewed articles | Other publications | Selected presentations

Barres B, B Montague-Hellen, and JB Yoder. 2017. Coming out: the experience of LGBT+ people in STEM. Genome Biology. 18:62 doi: 10.1186/s13059-017-1198-y. Invited Q and A panel for a series on diversity in science.

Yoder JB and J Leebens-Mack. 2016. The evolutionary ecology of “mutual services” in the twenty-first century. American Journal of Botany 103(10): 1712-16. doi: 10.3732/ajb.1600367. Full text (PDF, 623KB). Introduction to a Special Section on the ecology, genetics, and coevolution of intimate mutualisms.

Yoder JB. 2016. Coevolution, Introduction to. In: Kliman RM (ed.), Encyclopedia of Evolutionary Biology. Vol. 1: 314–21. Oxford: Academic Press. doi: 10.1016/B978-0-12-800049-6.00185-2.

Yoder JB. 2008. Science 2.0: Not so New? Science 320(5881): 1290. [with response by B. Shneiderman under the same title] doi: 10.1126/science.320.5881.1290. Full text (PDF, 521KB).

Selected presentations

Top | Peer-reviewed articles | Other publications | Selected presentations

Floral symmetry and specialization in pollination networks. Submitted presentation, American Society of Naturalists biennial meeting, 4-6 Jan 2020, Asilomar, California.

Species interactions and the origins of biodiversity. Invited presentation at the University of Texas Arlington, 26 Sept 2019, Arlington, Texas.

Floral symmetry and the structure of pollination networks. Presentation in a Spotlight Session organized for the American Society of Naturalists, at the Evolution meetings, 21-25 June 2019, Providence, Rhode Island.

Evolution and diversification through the lens of local adaptation. Invited plenary presentation in the Ecology and Evolutionary Biology Graduate Student Symposium at Yale University, 12 Apr. 2019, New Haven, Connecticut.

Queer in STEM: A study of sexualities and gender diversity in science, technology, engineering, and mathematics. Invited keynote presentation in the Queer in STEM Symposium, Yale University, 12 Apr. 2019, New Haven, Connecticut. Co-presented with D Cruz Ramirez de Arellano and J Nelson.

LGBTQ+ in STEM Demographics. Invited plenary presentation at the 2019 Out to Innovate conference, 17 Mar. 2019, Los Angeles, California. Co-presented with A Mattheis.

Local adaptation on complex landscapes. Invited presentation in the 2018 BMCB-GGD Symposium at Cornell University, 9 Oct. 2018, Ithaca, New York.

Finding local adaptation on complex landscapes. Invited presentation at California State University Los Angeles, 20 Sept. 2018, Los Angeles, California.

Mutualism, coevolution, and the origins of biodiversity. Invited presentation at California State University Dominguez Hills, 19 Apr. 2018, Los Angeles, California.

An introduction to current research on LGBTQ+ identities in STEM. Invited discussant in a symposium session organized by the National Organization of Gay and Lesbian Scientists and Technical Professionals at the AAAS annual meeting, 18 Feb. 2018, Austin, Texas.

Sanctions, partner recognition, and variation in mutualism. Submitted presentation at the American Society of Naturalists biennial meeting, 6 Jan. 2018, Pacific Grove, California.

Mutualism, coevolution, and the origins of biodiversity. Invited presentation at the University of California, Santa Barbara, 27 Nov. 2017, Santa Barbara, California.

Mutualism, coevolution, and the origins of biodiversity. Invited presentation at the University of California, Riverside, 26 Oct. 2017, Riverside, California.

Mutualism, coevolution, and the origins of biodiversity. Invited presentation at Rancho Santa Ana Botanic Garden, 22 Sept. 2017, Claremont, California.

Parallelized local adaptation in lodgepole pine (listed as Local adaptation and adaptive variation in distantly-related conifers). Submitted presentation at Evolution 2017 meetings, 23-27 June 2017, Portland, Oregon.

Sanctions, partner recognition, and variation in mutualism. Invited presentation at the Canadian Society for Ecology & Evolution Meeting 2017, 7-11 May 2017, Victoria, British Columbia.

Ecological genomics of parallel adaptation to climate in lodgepole pine and interior spruce. Submitted presentation at Evolution 2016 meetings, 18-21 June 2016, Austin, Texas. Video on YouTube.

Sanctions, partner recognition, and variation in mutualistic symbiosis. Invited presentation at EVO-WIBO, 16 Apr. 2016, Port Townsend, Washington. Winner, postdoctoral presentation award.

Using genomic data to identify and understand locally adapted genes. Invited presentation at University of Arizona, 5 Apr. 2016, Tucson, Arizona.

Dynamics and population genomics of mutualistic symbiosis. Presented in Colloquium: Integrated perspectives on the ecology, genetics and coevolution of intimate mutualisms, co-chaired with J Leebens-Mack. Botany 2015 meetings, 26 to 30 July 2015, Edmonton, Alberta.

Linking genomic pattern to evolutionary process. Invited presentation in Museum of Natural Science Seminar Series, Louisiana State University. 24 Oct. 2014, Baton Rouge, Louisiana.

Population genetic and genomic identification of locally adapted loci. Submitted presentation at Evolution meetings, 20 to 24 June 2014, in Raleigh, North Carolina. Talk slides (PDF, 2.29MB); video on YouTube.